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  1. Hamelin, F. M., Bowen, B., Bernhard, P., & Bokil, V. A. (2021). Optimal Control of Plant Disease Epidemics with Clean Seed Usage. Bulletin of Mathematical Biology, 83(5), 1-24. pdf

  2. Clin, P., Grognard, F., Mailleret, L., Val, F., Andrivon, D., & Hamelin, F.M. (2020). Taking advantage of pathogen diversity and immune priming to minimize disease prevalence in host mixtures: a model. Phytopathology, (ja). pdf

  3. Hilker, F. M., Sun, T. A., Allen, L. J. S., & Hamelin, F. M. (2020). Separate seasons of infection and reproduction can lead to multi-year population cycles. Journal of Theoretical Biology, 489, 110158. pdf

  4. Hamelin, F. M., Allen, L. J., Bokil, V. A., Gross, L. J., Hilker, F. M., Jeger, M. J., ... & Cunniffe, N. J. (2019). Coinfections by noninteracting pathogens are not independent and require new tests of interaction. PLoS biology, 17(12), e3000551. pdf

  5. Allen, L. J., Bokil, V. A., Cunniffe, N. J., Hamelin, F. M., Hilker, F. M., & Jeger, M. J. (2019). Modelling Vector Transmission and Epidemiology of Co-Infecting Plant Viruses. Viruses, 11(12), 1153. pdf

  6. Leclerc, M., Clément, J. A., Andrivon, D., & Hamelin, F. M. (2019). Assessing the effects of quantitative host resistance on the life-history traits of sporulating parasites with growing lesions. Proceedings of the Royal Society B, 286(1912), 20191244. pdf

  7. Calcagno, V., Hamelin, F., Mailleret, L., & Grognard, F. (2019). How optimal foragers should respond to habitat changes: on the consequences of habitat conversion. Theoretical Ecology, 1-11. pdf

  8. Hamelin, F. M., Hilker, F. M., Sun, T. A., Jeger, M. J., Hajimorad, M. R., Allen, L. J., & Prendeville, H. R. (2017). The evolution of parasitic and mutualistic plant–virus symbioses through transmission-virulence trade-offs. Virus Research, 241, 77-87. pdf

  9. Hilker, F. M., Allen, L. J., Bokil, V. A., Briggs, C. J., Feng, Z., Garrett, K. A., Gross, L. J., Hamelin, F. M., Jeger, M. J., Manore, C. A., Power, A. G., Redinbaugh, M. G., Rua, M. A., & Cunniffe, N. J. (2017). Modeling virus coinfection to inform management of maize lethal necrosis in Kenya. Phytopathology, 107(10), 1095-1108. pdf

  10. Ravigné, V., Lemesle, V., Walter, A., Mailleret, L., & Hamelin, F. M. (2017). Mate limitation in fungal plant parasites can lead to cyclic epidemics in perennial host populations. Bulletin of Mathematical Biology, 79(3), 430-447. pdf

  11. Hamelin, F. M., Bisson, A., Desprez-Loustau, M. L., Fabre, F., & Mailleret, L. (2016). Temporal niche differentiation of parasites sharing the same plant host: oak powdery mildew as a case study. Ecosphere, 7(11). pdf

  12. Hamelin, F. M., Castella, F., Doli, V., Marçais, B., Ravigné, V., & Lewis, M.A. (2016). Mate finding, sexual spore production, and the spread of fungal plant parasites. Bulletin of Mathematical Biology, 78(4), 695-712. pdf

  13. Fournet, S., Eoche-Bosy, D., Renault, L., Hamelin, F. M., & Montarry, J. (2016). Adaptation to resistant hosts increases fitness on susceptible hosts in the plant parasitic nematode Globodera pallida. Ecology and Evolution, 6(8), 2559-2568. pdf

  14. Hamelin, F. M., Allen, L. J., Prendeville, H. R., Hajimorad, M. R., & Jeger, M. J. (2016). The evolution of plant virus transmission pathways. Journal of Theoretical Biology, 396, 75-89. pdf

  15. Bernhard, P., & Hamelin, F. (2016). Sharing a resource with randomly arriving foragers. Mathematical Biosciences, 273, 91-101. pdf

  16. Soubeyrand, S., Sache, I., Hamelin, F., & Klein, E. K. (2015). Evolution of dispersal in asexual populations: to be independent, clumped or grouped? Evolutionary Ecology, 29(6), 947-963. pdf

  17. Calcagno, V., Grognard, F., Hamelin, F. M., Wajnberg, E., & Mailleret, L. (2014). The functional response predicts the effect of resouCasrce distribution on the optimal movement rate of consumers. Ecology Letters, 17(12), 1570-1579. pdf

  18. Bernhard, P., & Hamelin, F. M. (2014). Simple signaling games of sexual selection (Grafen’s revisited). Journal of Mathematical Biology, 69(6-7), 1719-1742. pdf

  19. Castel, M., Mailleret, L., Andrivon, D., Ravigné, V., & Hamelin, F. M. (2014). Allee effects and the evolution of polymorphism in cyclic parthenogens. The American Naturalist, 183(3), E75-E88. pdf

  20. Andrivon, D., Montarry, J., Corbière, R., Pasco, C., Glais, I., Marquer, B., Clément, J.A.J., Castel, M., & Hamelin, F. M. (2013). The hard life of Phytophthora infestans: when trade-offs shape evolution in a biotrophic plant pathogen. Plant Pathology, 62, 28-35. pdf

  21. Mailleret, L., Castel, M., Montarry, J., & Hamelin, F. M. (2012). From elaborate to compact seasonal plant epidemic models and back: is competitive exclusion in the details? Theoretical Ecology, 5(3), 311-324. pdf

  22. Hamelin, F. M., Castel, M., Poggi, S., Andrivon, D., & Mailleret, L. (2011). Seasonality and the evolutionary divergence of plant parasites. Ecology, 92(12), 2159-2166. pdf

  23. Montarry, J., Hamelin, F. M., Glais, I., Corbière, R., & Andrivon, D. (2010). Fitness costs associated with unnecessary virulence factors and life history traits: evolutionary insights from the potato late blight pathogen Phytophthora infestans. BMC Evolutionary Biology, 10(1), 283. pdf

  24. Hamelin, F. M., & Lewis, M. A. (2010). A differential game theoretical analysis of mechanistic models for territoriality. Journal of Mathematical Biology, 61(5), 665-694. pdf

  25. Hamelin, F., & Bernhard, P. (2008). Uncoupling Isaacs equations in two-player nonzero-sum differential games. Parental conflict over care as an example. Automatica, 44(3), 882-885. pdf

  26. Hamelin, F., Bernhard, P., Shaiju, A. J., & Wajnberg, E. (2007). Diet selection as a differential foraging game. SIAM journal on Control and Optimization, 46(5), 1539-1561. pdf

  27. Hamelin, F., Bernhard, P., & Wajnberg, É. (2007). Superparasitism as a differential game. Theoretical Population Biology, 72(3), 366-378. pdf

  28. Wajnberg, E., Bernhard, P., Hamelin, F., & Boivin, G. (2006). Optimal patch time allocation for time-limited foragers. Behavioral Ecology and Sociobiology, 60(1), 1-10. pdf


  1. Desprez-Loustau, M.-L., Hamelin, F.M., & Marçais, B. (2019). Understanding the recent ecological and evolutionary trajectory of a forest pathosystem: oak powdery mildew one century after the invasion in Europe. In Wildlife Disease Ecology: Linking Theory to Data and Application (Chap. 15), Wilson, K., Fenton, A., Tompkins, D. (Eds.). Cambridge University Press. pdf

  2. Bernhard, P., & Hamelin, F. (2009). Two-by-two static, evolutionary, and dynamic games. In From Semantics to Computer Science : Essays in Honour of Gilles Kahn (pp 465-487), Bertot, Y., Huet, G., Lévy J.-J., & Plotkin, G. (eds.). Cambridge University Press. pdf

  3. Hamelin, F., Bernhard, P., Shaiju, A.J., & Wajnberg, É. (2007). Foraging Under Competition: Evolutionarily Stable Patch-Leaving Strategies with Random Arrival Times. 2. Interference competition. In Advances in Dynamic Game Theory (pp. 349-365), Jørgensen S., Quincampoix, M., & Vincent T.L. (Eds.). Birkhäuser, Boston. pdf

  4. Hamelin, F., Bernhard, P., Nain, P., & Wajnberg, É. (2007). Foraging Under Competition: Evolutionarily Stable Patch-Leaving Strategies with Random Arrival Times. 1. Scramble competition. In Advances in Dynamic Game Theory (pp. 327-348), Jørgensen, S., Quincampoix, M., & Vincent, T.L. (Eds.). Birkhäuser, Boston. pdf

In French

  1. Tellier, A., & Hamelin, F.M. (2021) Coévolution des hôtes et de leurs parasites : la théorie. L’immunité végétale. Editions Quae.

  2. Bacaër, N., Hamelin, F.M., & Inaba, H. (2021). De nouvelles propriétés du pic épidémique. Quadrature, 119. pdf

PhD thesis

  • Hamelin, F. (2007). Jeux dynamiques en écologie du comportement. Doctoral dissertation, Université Nice Sophia Antipolis. pdf